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Any of them that look brown/maroon/have stress marks. Or any that you are thinking of disturbing. It is good to have a spare on hand if things don't cooperate. This can also include the radiator fittings themself.
Buy a can of silicone spray or rubber care aerosol and spray that into the female end of the couplings before prying on them. Use the same product to lube the fittings and orings before inserting.
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